Growth, development and nutritional manipulation of marbling in cattle: a review

By D. Pethick, G. Harper, V. Oddy. 2004

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Intramuscular fat content in meat-producing animals: development, genetic and nutritional control, and identification of putative markers. (2010, background): “Diets with a high glycaemic index (i.e. allowing rapid glucose absorption and concomitant high insulin levels) will deliver increased levels of net energy for lipogenesis; this explains why grain feeding promotes more IMF development compared with grass finishing (Pethick et al., 2004).”
Innovations in beef production systems that enhance the nutritional and health value of beef lipids and their relationship with meat quality. (2006, background): “At the cellular level, this may be due to increased levels of anabolic hormones (insulin) which stimulates lipogenesis and/or to a preference of marbling adipocytes for carbohydrate carbon to synthesise fatty acids unlike adipocytes of the carcass (Pethick et al., 2004).”