Ectopic expression of nicotianamine synthase genes results in improved iron accumulation and increased nickel tolerance in transgenic tobacco

By D. Douchkov, C. Gryczka, U. W. Stephan, R. Hell, H. Bäumlein. 2005

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Biofortification of crops for reducing malnutrition (2012, background): “Nicotianamine (NA), a chelator of metals, is ubiquitous in higher plants and a key ligand involved in metal transport and homeostasis (Douchkov et al. 2005).”
Molecular mechanisms of metal hyperaccumulation in plants. (2009, background): “Increase in Ni tolerance could be gained upon NAS overexpression in nontolerant species (Douchkov et al., 2005; Kim et al., 2005; Pianelli et al., 2005).”
Functions and toxicity of nickel in plants: recent advances and future prospects. (2009, background): “[78] D.”
Biofortification of crops with seven mineral elements often lacking in human diets--iron, zinc, copper, calcium, magnesium, selenium and iodine. (2009, background): “, 2004; Grotz & Guerinot, 2006), and in strategy II plants by increasing the synthesis and exudation of phytosiderophores (Takahashi et al., 2001; Douchkov et al., 2005), together with increased expression of genes encoding YSL proteins.”