Lignification: are lignins biosynthesized via simple combinatorial chemistry or via proteinaceous control and template replication?

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The genetics of lignin biosynthesis: connecting genotype to phenotype. (2010, background): “The arguments and counterarguments that have been put forth by the proponents of these two views are far too numerous to list here and have been thoroughly treated elsewhere (19, 20, 43, 71, 91, 95, 105).”
The cell biology of lignification in higher plants. (2015, background): “Both the polymer length and the proportion of different linkages can be explained by the availability and reactivity of specific lignin monomer(s) radicals (Ralph et al., 2008).”
Phenolic Compounds: Introduction 50 (2013, background): “5 Spruce lignin model (Redrawn from [87]) 50 Phenolic Compounds: Introduction 1563”
Cell wall deposition and metabolism in wheat grain (2012, background): “How monolignols are polymerized is still open to debate (Davin et al., 2008; Ralph et al., 2008) but lignin structure is clearly influenced by the polysaccharidic matrix (Aimi et al.”
Lignocellulosic biomass: Biosynthesis, degradation, and industrial utilization (2016, background): “However, further concrete mechanistic evidence is needed to displace the currently held view that the polymerization step in lignans and/or lignins is entirely under chemical control [113].”