Distribution, biosynthesis and function of purine and pyridine alkaloids in Coffea arabica seedlings

By Xinqiang Zheng, H. Ashihara. 2004

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Cytology, biochemistry and molecular changes during coffee fruit development (2006, background): “In coffee plants, caffeine is actively biosynthesized during leaflet emergence and then decreases when leaves reach their optimal photosynthetic capacity (Frischknecht et al., 1986; Ashihara et al., 1996; Zheng and Ashihara, 2004).”