Ambiguous role of phlorotannins as chemical defenses in the brown alga Fucus vesiculosus

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The genus Laminaria sensu lato : recent insights and developments (2008, background): “Similarly, phlorotannin-enriched feed did not adversely affect amphipod fitness (Kubanek et al , 2004 ).”
Red and green macroalgae for fish and animal feed and human functional food development (2016, background): “Also, several studies have linked the presence of phlorotannins and other associated metabolites, such as sulfated sugars, as a chemical defense mechanism against herbivores in brown seaweeds.((116)) Macroalgae may also contain variable amounts of nitrogen as ammonium instead of protein.”