Inorganic polyphosphate is needed for swimming, swarming, and twitching motilities of Pseudomonas aeruginosa.

By M. H. Rashid, A. Kornberg. Proceedings of the National Academy of Sciences of the United States of America, 2000

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Negative Control of Quorum Sensing by RpoN (σ54) in Pseudomonas aeruginosa PAO1 (2003, methodology): “Swarming was tested on plates containing 0.5% (wt/vol) agar, 8 g of nutrient broth per liter (Oxoid), 5 g of glucose per liter, and 1 mM L-glutamine (51).”
Pseudomonas aeruginosa population structure revisited under environmental focus (2010, background): “Twitching is necessary for infectivity and biofilm formation process (Beatson et al., 2002a; Beatson et al., 2002b; Fraser & Hughes, 1999; Leone I., 2008; Rashid & Kornberg, 2000).”
Swarming of Pseudomonas aeruginosa Is Dependent on Cell-to-Cell Signaling and Requires Flagella and Pili (2000, background): “During preparation of the manuscript, an article by Rashid and Kornberg (44) also reported on swarming motility in P. aeruginosa.”
A field guide to bacterial swarming motility (2010, neutral): “aeruginosa , which swims with a single polar flagellum, may produce two polar flagella when moving on a surfac”
Cell Death in Pseudomonas aeruginosa Biofilm Development (2003, neutral): “3% agar, respectively, as described elsewhere (49).”
Attachment to and biofilm formation on abiotic surfaces by Acinetobacter baumannii: involvement of a novel chaperone-usher pili assembly system. (2003, methodology): “Swimming, swarming, sliding and twitching agar plates (Rashid & Kornberg, 2000) were used to test different types of cell motility.”