Glutamate is an inhibitory neurotransmitter in the Drosophila olfactory system

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Dissecting neural pathways for forgetting in Drosophila olfactory aversive memory (2015, background): “Here, the receptor signaling (75, 76) activated by MBON-γ4>γ1γ2 remains to be defined.”
Heterosynaptic plasticity underlies aversive olfactory learning in Drosophila (2015, unknown): “Since the cell body of MBON-g2a01 was located ventral to the antennal lobes, we had to rotate the head by 180 at the neck to expose the ventral side of the brain for recording, as described previously (Liu and Wilson, 2013).”
Activity of Defined Mushroom Body Output Neurons Underlies Learned Olfactory Behavior in Drosophila (2015, background): “, 2013) neurons, whereas the horizontal outputs are onto glutamatergic, potentially inhibitory (Liu and Wilson, 2013), M4b/MBON-b2b02a, M4b0/MBON-b02mp, and M6/MBON-g5b02a neurons.”
The Drosophila Mushroom Body: From Architecture to Algorithm in a Learning Circuit. (2020, background): “In the MB, there is a single large inhibitory neuron, termed APL, that innervates the entirety of the MB lobes and calyx (Liu & Davis 2009).”
The complete connectome of a learning and memory centre in an insect brain (2017, background): “Glutamate has previously been found to be inhibitory in the ﬂy 46 , but an excitatory effect on some neurons cannot be excluded.”