Metal-specific and NADPH oxidase dependent changes in lipoxygenase and NADPH oxidase gene expression in Arabidopsis thaliana exposed to cadmium or excess copper.

By Tony Remans, Kelly Opdenakker, K. Smeets, Dennis Mathijsen, Jaco Vangronsveld, A. Cuypers. 2010

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The molecular mechanism of zinc and cadmium stress response in plants (2012, background): “NADPH oxidase is localized on the plasma membrane and produces superoxide (O2 ) from oxygen (O2) when challenged with excess metals [176– 178].”
Molecular Mechanism of Heavy Metal Toxicity and Tolerance in Plants: Central Role of Glutathione in Detoxification of Reactive Oxygen Species and Methylglyoxal and in Heavy Metal Chelation (2012, background): “NADPH oxidase-dependent ROS induction has been reported in wheat in response to Ni stress [221], in pea (Pisum sativum) in response to Cd stress [222], in Vicia faba in response to Pb stress [223], and in Arabidopsis in response to Cd and Cu [224].”
Glutathione Is a Key Player in Metal-Induced Oxidative Stress Defenses (2012, background): “Finally, metals can activate pro-oxidative enzymes such as NADPH oxidases and lipoxygenases [13,93,98,99].”