Tracking biological invasions in space and time: elucidating the invasive history of the green alga Codium fragile using old DNA

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Introduced and cryptogenic marine and estuarine species of South Africa (2011, background): “Provan et al. (2008) stated that this alga had recently been reported from South Africa “in 1999”, citing Begin and Scheibling (2003).”
European seaweeds under pressure: Consequences for communities and ecosystem functioning (2015, background): “fragile (Provan et al., 2008), pinpointing the North West Pacific as the native area.”
GENETIC DATA HINT AT A COMMON DONOR REGION FOR INVASIVE ATLANTIC AND PACIFIC POPULATIONS OF GRACILARIA VERMICULOPHYLLA (GRACILARIALES, RHODOPHYTA) 1 (2010, result): “The low genetic variation in introduced G. vermiculophylla is consistent with previous reports on other introduced seaweeds, including Caulerpa taxifolia (Jousson et al. 2000) and Codium fragile subsp. tomentosoides (Provan et al. 2005, 2008).”
Temperature-mediated outbreak dynamics of the invasive bryozoan Membranipora membranacea in Nova Scotian kelp beds (2009, background): “…an epiphytic bryozoan Membranipora membranacea and an invasive green alga Codium fragile ssp. fragile (formerly C. fragile ssp. tomentosoides; Provan et al. 2008; hereafter C. fragile) resulted in displacement of native kelps throughout this region (Chapman et al. 2002, Levin et al. 2002,…”