Flexible foraging techniques in breeding Cormorants Phalacrocorax carbo and Shags Phalacrocorax aristotelis: benthic or pelagic feeding?

By D. Grémillet, G. Argentin, B. Schulte, B. Culik. 2008

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Microhabitat use and prey capture of a bottom-feeding top predator, the European shag, shown by camera loggers (2008, background): “1991b, Watanuki et al. 2005, but see Grémillet et al. 1998, Wanless et al. 1998) on lesser sandeels Ammodytes marinus (Harris & Wanless 1991).”
Confusion reigns? A review of marine megafauna interactions with tidal-stream environments (2015, background): “However, there are also distinct dietary variations at species-level, even within geographically similar areas, both between individuals (Elliott et al. 2008) and for the same individuals over time (Grémillet et al. 1998).”