Co-expression of the Suaeda salsa SsNHX1 and Arabidopsis AVP1 confer greater salt tolerance to transgenic rice than the single SsNHX1

By F. Zhao, Xue-Jie Zhang, Pinghua Li, Yanxiu Zhao, Hui Zhang. Molecular breeding, 2006

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Engineering Salinity and Water-Stress Tolerance in Crop Plants (2011, background): “Transformation of rice with the SOD2 gene (under 35S promoter) resulted in accumulation of more Kþ, Ca2þ, Mg2þ and less Naþ in the shoots compared with wild type (Zhao et al., 2006b).”
Overexpression of the Arabidopsis H+-PPase enhanced resistance to salt and drought stress in transgenic alfalfa (Medicago sativa L.). (2009, background): “This is consistent with the phenotype observed from transgenic rice co-expressing the SsNHX1 and AVP1 [14].”
Overexpression of OsVP1 and OsNHX1 Increases Tolerance to Drought and Salinity in Rice (2010, background): “The electrochemical potential gradient of H between the cytoplasm and vacuole can promote the exchange of Na/H through antiporters and enable the transport of some small organic acids and sugars (Zhao et al. 2006).”
Smart Engineering of Genetic Resources for Enhanced Salinity Tolerance in Crop Plants (2016, background): “In rice, co-expression of SsNHX1 and AtAVP1 in transgenic plants conferred higher salinity tolerance than that of the individual SsNHX1 (Zhao et al., 2006).”