Cockroach aggregation based on strain odour recognition

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The principles of collective animal behaviour (2006, background): “There is evidence that only ant species with large colonies use pheromone trails for communicating the presence of food (Beckers et al. 1989), but there is little evidence from between species comparison that individual complexity decreases with increased colony size (Anderson & McShea 2001).”
A review of swarm robotics tasks (2016, background): “The aggregation behavior of cockroaches is similar to that of honeybees, but cockroaches have in addition the ability to detect the number of neighbors, and the probabilities of entering the wait and walk states are dependent on the number of detected neighbors [51].”
Segregation of Heterogeneous Units in a Swarm of Robotic Agents (2010, background): “Several species of cockroaches aggregate or segregate [1] based on odors of strains.”
Self-organized aggregation in cockroaches (2005, background): “In both cases, the probability of a cockroach leaving a resting site decreased as a function of the number of conspecifics in the site (Rivault & Cloarec 1998; Amé et al. 2004).”