New examples of holococcolith¯heterococcolith combination coccospheres and their implications for coccolithophorid biology.

By L. Cros, A. Kleijne, A. Zeltner, C. Billard, J. Young. 2000

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A review of selected aspects of coccolithophore biology with implications for paleobiodiversity estimation (2005, background): “In addition, however, combination coccospheres bearing coccoliths typical of two phases of the life-cycle occasionally occur and observation of these has greatly supplemented our knowledge of life cycles (e.g. Cros et al. 2000; Geisen et al. 2002).”
The Biodiversity of the Mediterranean Sea: Estimates, Patterns, and Threats (2010, background): “lists 166 species of coccolithophorids of the northwestern Mediterranean and revised the classification of several important taxa [see also 160]. Recently, the discovery of a number of combination coccospheres bearing holo- and heterococcoliths”
Atlas of Northwestern Mediterranean Coccolithophores (2002, background): “A member of this family, Helicosphaera carteri, has been shown to form combination coccospheres with holococcoliths (Cros et al., 2000).”
What is new in coccolithophore biology (2004, background): “Polycrater galapagensis produces unusual aragonitic coccoliths (often considered as nannoliths) and its assignment to the haptophytes was tentative until it was shown to possess a hap- What is new in coccolithophore biology?”