Deformation mechanisms in nacre

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Sacrificial bonds and hidden length dissipate energy as mineralized fibrils separate during bone fracture (2005, background): “There may also be a significant mechanical effect of the roughness of the mineralized fibrils, especially in tension along the fibrils; such a nanoscale friction mechanism has been proposed for nacr”
Nanoscale Morphology and Indentation of Individual Nacre Tablets from the Gastropod Mollusc Trochus Niloticus (2005, background): “It has also been suggested that interacting nanoasperities and mechanical interlocking provide resistance to interfacial sliding, thus determining the level of stress needed to initiate inelastic strain.(3,37) The existence of a finer nanoasperity structure arranged into assemblies of larger nanoasp”
The conflicts between strength and toughness. (2011, background): “, the frictional motion between mineral platelets in seashell”
Biological materials: Functional adaptations and bioinspired designs (2012, background): “The mechanical properties of nacre were first studied by Currey [61,62,352–355] followed by Jackson [55,356,357], Heuer [50,358], Sarikaya and Aksay [37,318,324,359–361], Meyers [67–69,348] and Evans [34,218,350].”
Biomimetics: lessons from nature–an overview (2009, background): “The deformation mechanisms and mechanics of nacre have been investigated by Wang et al. (2001) and Barthelat et al. (2007).”