Factors and scales potentially important for saproxylic beetles in temperate mixed oak forest

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Hollow oaks and beetle conservation: the significance of the surroundings (2010, background): “The importance of dead wood in the surroundings has proven influential in other studies of saproxylic beetles (Økland et al. 1996; Franc et al. 2007).”
Increased openness around retained oaks increases species richness of saproxylic beetles (2012, background): “According to other studies, both local, landscape and regional factors are important for local species richness of saproxylic oak-associated beetles in mixed oak-rich forests, such as the local quantity of dead oak wood (Götmark et al. 2011; Franc et al. 2007).”