Habitat heterogeneity, species diversity and null models

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Biodiversity along temperate forest succession (2018, background): “Another factor is the diversity of resources (Cramer & Willig, 2005).”
The influence of habitat integrity and physical-chemical water variables on the structure of aquatic and semi-aquatic Heteroptera (2010, background): “Species become more diverse with increasing environmental heterogeneity, resulting in increased niche partitioning, which optimizes specialization and decreases the effects of inter-specific competition (O’CONNOR 1991, CRAMER & WILLIG 2005).”
Do Neotropical peccary species (Tayassuidae) function as ecosystem engineers for anurans? (2010, methodology): “We chose the Morisita–Horn index because it is unbiased with respect to sample size and richness, and therefore is one of the most robust β-similarity indices (Cramer & Willig 2004, Magurran 2003).”