Plant cell membrane as a marker for light-dependent and light-independent herbicide mechanisms of action.

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Chlorophyll fluorescence as a marker for herbicide mechanisms of action (2012, background): “Pelargonic acid, or nonanoic acid (Table 4), induces light-independent membranotropic destabilization as well as a light-dependent membrane peroxidation from radicals derived from photosensitized chlorophyll displaced from the thylakoid membranes [24,63].”
Biochemical Markers and Enzyme Assays for Herbicide Mode of Action and Resistance Studies (2015, methodology): “Therefore, the simple electrolyte leakage experiment can easily identify the light-dependent nature of these herbicides (see ‘‘Electrolyte Leakage’’ in the ‘‘Common Protocols’’ section) (Dayan and Watson 2011).”