The role of chromatic and achromatic signals for fruit detection by birds

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Role of Anthocyanins in Plant Defence (2008, background): “It is likely, therefore, that both chromatic (wavelength related) and achromatic (intensity related) information is involved, as is known also to be used by frugivorous birds (Schaefer et al. 2006).”
Iridescent structurally based coloration of eyespots correlates with mating success in the peacock (2007, background): “Because both achromatic (intensity related) and chromatic (wavelength related) signals are important for object detection (e.g., for birds: Osorio, Miklosi, and Gonda 1999; Osorio, Vorobyev, and Jones 1999; Schaefer et al. 2006), it is important to analyze colors using a physiological model of visio”
Why are fruits colorful? The relative importance of achromatic and chromatic contrasts for detection by birds (2009, background): “First, in the lone experiment on this subject Schaefer et al. (2006) found that—similar to our results—crows prioritized chromatic over achromatic contrasts when searching for fruits among foliage.”
Birds use fruit colour as honest signal of dietary antioxidant rewards (2008, unknown): “…conspicuousness appears counterintuitive, but it is supported by an experiment documenting that some birds can detect red fruits that have typically low contents of anthocyanins from a larger distance than black fruits that are characterised by high contents of anthocyanins (Schaefer et al. 2006).”