Functional association of bill morphology and foraging behaviour in calidrid sandpipers

By S. Nebel, D. Jackson, R. Elner. 2005

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The relative importance of olfaction and remote touch in prey detection by North Island brown kiwis (2009, background): “‘Remote touch’ is mediated by an organ composed of numerous pits in the bone of the bill tips packed with Herbst corpuscles and terminal cell receptors (Grandry corpuscles; Bolze 1968; Piersma et al. 1998; Nebel et al. 2005).”
Visual fields and their functions in birds (2007, background): “These include species that locate prey using tactile cues derived from sensory pits containing Herbst and Grandry corpuscles located around the tips of the maxilla and mandible (Gottschaldt 1985; Nebel et al. 2005; Piersma et al. 1998), as are found in long-billed probing shorebirds, for example Woo”