Swarming in honey bees (Apis mellifera) and Varroa destructor population development in Sweden.

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Characteristics of honey bee colonies (Apis mellifera) in Sweden surviving Varroa destructor infestation (2011, background): “For a more detailed explanation on the background of these surviving honey bee colonies, refer to Fries et al. (2003; 2006) and Fries and Bommarco (2007).”
Biology and control of Varroa destructor. (2010, background): “…colony as a ‘‘superorganism”, is reduced in two ways, even if the infestation is moderate: Drones which have been parasitized during their development have a significantly lower chance to mate (Duay et al., 2002) and infested colonies produce less swarms (Fries et al., 2003; Villa et al., 2008).”
The influence of genetic origin and its interaction with environmental effects on the survival of Apis mellifera L. colonies in Europe (2014, background): “Indeed, swarming has been shown to have an effect on mite infestation levels and, consequently, on colony survival (Fries et al., 2003).”
Occurrence of parasites and pathogens in honey bee colonies used in a European genotype-environment interactions experiment (2014, background): “Swarming affects V. destructor reproduction due to the interruption of bee brood production, although the results of Fries et al. (2003) demonstrated that swarming did not have a significant effect on the build up of detrimental mite levels.”