Is phytoplankton growth in the Wadden Sea light or nitrogen limited

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Spatial variation in phytoplankton dynamics in the Belgian coastal zone of the North Sea studied by microscopy, HPLC-CHEMTAX and underway fluorescence recordings (2006, background): “In the shallow, turbid waters of the southern North Sea, light is an important factor regulating phytoplankton development (Gieskes and Kraay, 1975; Tett and Walne, 1995; Colijn and Cadée, 2003).”
Effects of climate change on growth of 0-group sole and plaice (2008, background): “As the primary production in coastal waters such as the Wadden Sea is mainly determined by light and nutrients (Colijn & Cadée 2003), an increase in summer temperature due to global warming may raise the food requirements of the secondary and tertiary consumers above primary production and result…”
Harmful algal blooms: How strong is the evidence that nutrient ratios and forms influence their occurrence? (2012, background): “Indeed, Colijn and Cadée (2003) were of the opinion that in the Wadden Sea and the Marsdiep region in particular, irradiance was more limiting for the production of phytoplankton biomass and there was only slight nutrient limitation between May and July.”