Why are small seeds dispersed through animal guts: large numbers or seed size per se?

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The functional role of soil seed banks in natural communities. (2014, background): “001 (A. Saatkamp, 2009, unpublished data, and data from Flynn et al., 2004; Holmes and Newton, 2004; Bruun and Poschlod, 2006; Soons et al., 2008; Gardarin et al., 2010; Morozowska et al., 2011).”
The LEDA Traitbase: a database of life‐history traits of the Northwest European flora (2008, background): “Seed number, seed shedding Response to disturbance, establishment, dispersal (Leishman 2001; Bruun & Poschlod 2006 )”
New handbook for standardised measurement of plant functional traits worldwide (2013, background): “References on theory and significance: Bruun and Poschlod (2006); Poschlod et al. (1998, 2005) Tackenberg (2003); Tackenberg et al. (2003); Schurr et al.”
Seed dispersal by ungulates as an ecological filter: a trait-based meta-analysis (2015, background): “However, higher numbers of small seeds in dung samples do not necessarily appear to be related to a morphological advantage, but rather to their higher number in the vegetation (Bruun and Poschlod 2006).”