How important is natural behaviour in animal farming systems

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A review of behavioural methods to study emotion and mood in pigs, Sus scrofa (2014, background): “While no study has looked at physiological correlates of QBA ratings in pigs, Stockman et al. (2011) demonstrated that cattle rated as more “agitated” by observers also had higher heart rate and body temperature, among other measures.”
Invited review: Effects of heat stress on dairy cattle welfare. (2017, background): “In contrast, others argue that this interpretation and application of natural living into management practices may negatively affect welfare (e.g., by exposing the animal to diseases, parasites, extreme weather, and predators; Špinka, 2006).”
The importance of social behaviour for goat welfare in livestock farming (2010, background): “Evidently, 22 allowing and encouraging the animals to perform behaviours that are associated with 23 positive emotions improves their welfare (Špinka, 2006).”
Animal welfare with and without consciousness (2017, background): “…of ‘natural behaviour’: differences between the behaviour of captive and wild animals can be documented but it is then difficult to argue that the captive animal, safe from predation, has reduced welfare because it lacks the ‘natural’ behaviour of running away from a predator ( Spinka, 2006).”