Juvenile survival and benefits of play behaviour in brown bears, Ursus arctos

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Partner preferences and asymmetries in social play among domestic dog, Canis lupus familiaris, littermates (2008, unknown): “Only one study has linked play directly to survival (brown bear cubs, Ursus arctos, that play more in their first summer are more likely to survive until the end of the next summer), but the mechanisms involved remain unknown (Fagen & Fagen 2004).”
Play, Playfulness, Creativity and Innovation (2013, background): “A remarkable example of innovation is the cooperative foraging in small groups of hump-backed whales observed in Southeast Alaska and the west coast of South America.”
Animal play and animal welfare (2011, background): “…social play, and also nonsocial play in other species, increasingly suggest that play may indeed equip animals with the general-purpose competencies and resilience outlined above (Donaldson et al. 2002; Von Frijtag et al. 2002; Fagen & Fagen 2004; Dudink et al. 2006; see also Pellis & Pellis 2009).”
Why do Dolphins Play (2014, background): “…significance of play rests at least in part on the experiences it provides for players to learn to produce appropriate emotional responses in a variety of contexts (Bekoff, 2002; Colvin & Tissier, 1985; Fagen & Fagen, 2004; Kuczaj & Horback, 2013; Pellis & Pellis, 2009; Singer & Singer, 1990).”