History or abiotic filter: which is more important in determining the species composition of calcareous grasslands?

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Testing a relict distributional pattern of fen plant and terrestrial snail species at the Holocene scale: a null model approach (2011, background): “Although some studies have identified indicator species of either ancient grasslands (Karlı́k & Poschlod, 2009) or ancient forests (Honnay et al., 1998), these studies dealt with a timescale of a few hundred years.”