Residue allelopathy in Parthenium hysterophorus L.—Does parthenin play a leading role?

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Impact of Parthenium hysterophorus L. invasion on plant species composition and soil properties of grassland communities in Nepal (2011, background): “Additionally, these chemicals inhibit the growth of other plants through the contamination of soil (e.g., Batish et al., 2005; Belz et al., 2007; Kanchan, 1975; Picman and Picman, 1984; Rajan, 1973; Rashid et al., 2008; Singh et al., 2002, 2005).”
Synthesis, characterization and evaluation of effect of phytogenic zinc nanoparticles on soil exo-enzymes (2014, background): “Parthenin plays a pivotal role as root exudate in allelopathic interference with surrounding plants (Belz et al. 2007).”
Sesquiterpene Lactones: More Than Protective Plant Compounds With High Toxicity (2016, background): “Because STLs are released from the parent plant in nature mainly by two mechanisms, leaching by rain or decomposition of plant residues (Kanchan and Jayachandra 1980; Belz et al., 2007), phytotoxin concentrations under field conditions in leachates are commonly believed to be below inhibitory levels”