Facilitation of tree saplings by nurse plants: Microhabitat amelioration or protection against herbivores?

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Facilitation of Quercus ilex in Mediterranean shrubland is explained by both direct and indirect interactions mediated by herbs (2010, background): “…facilitation have focused on interactions between different trophic levels, such as those analysing defence against herbivory (Boulant et al. 2008; Gómez-Aparicio et al. 2008; Anthelme &Michalet 2009) or concentration of propagules of other species (Verdú & Garcı́a-Fayos 2003; Aerts et al.…”
A seeding experiment for testing tree-community recruitment under variable environments: Implications for forest regeneration and conservation in Mediterranean habitats (2009, background): “Most of these factors can fluctuate inter-annually due to variation in climatic conditions, thus altering the quality of the habitat for recruitment (e.g. Greenlee and Callaway, 1996; Tielbörger and Kadmon, 2000; Ibáñez and Schupp, 2001; Gómez-Aparicio et al., 2005, 2008).”
Shedding light on shade: ecological perspectives of understorey plant life (2016, background): “Interestingly, the relative importance of biotic and abiotic effects changes along stress gradients, and also depends on functional traits of the interacting species (Gómez-Aparicio et al. 2008).”