Niche partitioning between two sympatric genetically distinct cave bears (Ursus spelaeus and Ursus ingressus) and brown bear (Ursus arctos) from Austria: Isotopic evidence from fossil bones

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Late Quaternary megafaunal extinctions on the continents: a short review (2015, methodology): “In addition, stable isotope analyses are used to provide important information on the diets and community structure of Quaternary megafaunal species (e.g. Stevens and Hedges, 2004; Bocherens et al., 2011a, b).”
Isotopic evidence for dietary ecology of cave lion (Panthera spelaea) in North-Western Europe: Prey choice, competition and implications for extinction (2011, background): “Although cave bears belong to the order Carnivora, the isotopic data gathered for specimens from the studied sites clearly point to a vegetarian diet, ruling them out from dietary competition with lions (e.g., Bocherens et al., 1997; 2011; Münzel et al., 2008; Bocherens et al., in press).”