Yield and grain protein concentration in bread wheat: how to use the negative relationship between the two characters to identify favourable genotypes?

By F. Oury, C. Godin. Euphytica, 2007

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Breeding for increased nitrogen‐use efficiency: a review for wheat (T. aestivum L.) (2016, background): “…the cropping season and has a critical interaction with HI: once grains are growing, they become an N sink, and growers, breeders and the wheat industry have to manage the contradictory objectives of high yields and high protein contents (Feil 1997, Jeuffroy et al. 2002, Oury and Godin 2007).”
Relationships between grain protein content and grain yield components through quantitative trait locus analyses in a recombinant inbred line population derived from two elite durum wheat cultivars (2012, background): “The improvement of both traits has been hampered by the generally negative relationship between GPC and grain yield components in wheats, as well as in barley, maize, oat and sorghum (see review by Simmonds 1995; Feil 1997; Oury et al. 2003; Oury and Godin 2007).”