Clinal genetic variation and isolation by distance in the European eel Anguilla anguilla (L.)

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How does the European eel (Anguilla anguilla) retain its population structure during its larval migration across the North Atlantic Ocean (2006, background): “It is remarkable that the European eel (Anguilla anguilla) may retain regionally distinct population structure (e.g., Daemen et al. 2001; Wirth and Bernatchez 2001; Maes and Volckaert 2002) during a life cycle that incorporates two migrations across the North Atlantic Ocean at the beginning and end ”
Geographic effects on American eel (Anguilla rostrata) life history characteristics and strategies (2010, background): “More recent genetic analyses of anguillid eel nuclear DNA have focused on FST (the proportion of total genetic variation occurring among populations), with low but significant FST supporting the existence of weak genetic structure characterized by an isolation-by-distance (IBD) pattern for the Europ”
The European eel (Anguilla anguilla, Linnaeus), its Lifecycle, Evolution and Reproduction: A Literature Review (2005, background): “Finally, Maes and Volckaert (2002) reported clinal genetic structure and IBD in the European eel population using 15 allozyme loci and identified three distinct groups: Northern Europe, Western Europe and the Mediterranean Sea.”