From the Cover: The insect endosymbiont Sodalis glossinidius utilizes a type III secretion system for cell invasion

By C. Dale, S. Young, Daniel T. Haydon, S. Welburn. 2001

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Secondary (γ-Proteobacteria) Endosymbionts Infect the Primary (β-Proteobacteria) Endosymbionts of Mealybugs Multiple Times and Coevolve with Their Hosts (2002, background): “It has been shown that this organism requires a functional type III secretion system to establish itself in its insect host (14).”
The biology of blood-sucking in insects (2005, unknown): “For example, molecular approaches have shown that the tsetse endosymbiont Sodalis glossinidius utilizes a type III secretion system for cell invasion (Dale et al., 2001) suggesting that Sodalis may have evolved from an ancestor such as Salmonella with a parasitic intracellular lifestyle.”
Plants, mycorrhizal fungi, and bacteria: a network of interactions. (2009, background): “TTSS genes have been reported in animal secondary symbionts such as Sodalis glossinidius, “Candidatus Hamiltonella defensa” and Candidatus Arsenophonus arthropodicus” (29), whereas they were not detected in the genome of primary symbionts, such as Buchnera.”