Diversity in a complex ecological network with two interaction types

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Describe, understand and predict: why do we need networks in ecology? (2016, background): “As such multilayer data sets are also emerging recently in ecology (Meli an et al. 2009; Olesen et al. 2011; Pocock et al. 2012; K efi et al. 2015, 2016; Pilosof et al. 2015; Sander et al. 2015), the application of these new tools to ecological data sets will provide a more detailed understanding…”
The ‘night shift’: nocturnal pollen‐transport networks in a boreal pine forest (2011, background): “Studying networks that include more than one type of interaction is likely to improve our understanding of the architecture of biodiversity (Melián et al., 2008).”
The Structure of Ecological Networks Across Levels of Organization (2020, background): “In a network describing interactions between plants and their herbivores, pollinators, and seed dispersers, the ratio of mutualistic to antagonistic interactions varied widely across plants, with just a few plants showing very high mutualism to antagonism ratios (Melián et al. 2009).”
Structure–stability relationships in networks combining mutualistic and antagonistic interactions (2014, background): “Very few studies have considered community dynamics with both mutualistic and antagonistic interactions (but see, Ringel et al. 1996, Jang 2002, Allesina and Pascual 2008, Melián et al. 2009, Mougi and Kondoh 2012).”