Predicting the potential geographical distribution of the harlequin ladybird, Harmonia axyridis, using the CLIMEX model

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Quarantine arthropod invasions in Europe: the role of climate, hosts and propagule pressure (2014, background): “…Diptera), Halyomorpha halys St al (Niva & Takeda, 2003) (Pentatomidae; Hemiptera) Diabrotica virgifera (Baker et al., 2003), Leptinotarsa decemlineata Say (Kocm ankov a et al., 2011) (Chrysomelidae; Coleoptera) and Harmonia axyridis Pallas (Poutsma et al., 2008) (Coccinellidae; Coleoptera).”
The global spread of Harmonia axyridis (Coleoptera: Coccinellidae): distribution, dispersal and routes of invasion (2011, background): “Given the climatic conditions encountered in the native range, from tropical southern Japan to Siberia, it is likely that H. axyridis has a very broad temperature threshold.”
The harlequin ladybird, Harmonia axyridis: global perspectives on invasion history and ecology (2016, background): “axyridis (Poutsma et al. 2008), and in support of this there have only been occasional records of adults, arriving on produce imported from the mainland, from these northern islands (Ribbands et al.”