An energetic correlate between colony size and foraging effort in seabirds, an example of the Adélie penguinPygoscelis adeliae

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Top-down and bottom-up factors affecting seabird population trends in the California current system (1985-2006) (2010, background): “…and the overall productivity of the CCS, as well as the recovery of potentially major trophic competitors (whales), we test the prediction that speciesspecific energetic costs of prey acquisition influence the numerical responses of seabird species to food variability (Ballance et al., 2009).”
Foraging strategies of Adélie penguins: adjusting body condition to cope with environmental variability (2010, background): “If a single habitat is available, as in the present study, prey depletion may play an important role in determining foraging trip duration and, ultimately, foraging energetics, regardless of whether adults need to regain condition (see Ainley et al. 2004, 2006, Ballance et al. 2009).”
Central-Place Foraging in an Arctic Seabird Provides Evidence for Storer-Ashmole's Halo (2009, background): “In the absence of such manipulations, the abundance of indirect evidence from different taxa and locations supports Storer-Ashmole’s halo as an important determinant of variation in the breeding biology of a wide range of seabirds (Ainley et al. 2004, Gaston 2004, Ballance et al. 2009).”