Effects of shrub encroachment on herbage production and nutritive value in semi-arid Mediterranean grasslands

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Are shrubs really a sign of declining ecosystem function? Disentangling the myths and truths of woody encroachment in Australia (2014, background): “Similarly, foliage concentrations of fibre and lignin have been shown to increase with increasing shrub cover, although with no clear responses for organic matter digestibility (Zarovali et al. 2007).”
Impacts of shrub encroachment on ecosystem structure and functioning: towards a global synthesis (2011, background): “Compared with the original grasslands, the popular view is that shrublands are associated with reduced plant biomass and species richness, and a range of other effects on the vegetation (see below; Zarovalli et al. 2007; Van Auken 2009).”