Adaptive shifts in honey bee (Apis mellifera L.) guarding behavior support predictions of the acceptance threshold model

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The development of collective personality: the ontogenetic drivers of behavioral variation across groups (2014, background): “In honeybees , even though some colonies are consistently more aggressive than others (Pearce et al., 2001; Hunt, 2007), aggression levels increase with colony size (Wray et al., 2011), and/or when resources become scarce (Downs and Ratnieks, 2000).”
Wax combs mediate nestmate recognition by guard honeybees (2006, methodology): “We used a standard assay of guard discrimination (Downs & Ratnieks 2000).”
To fight or not to fight: context-dependent interspecific aggression in competing ants (2009, background): “For example, seasonal variation and/or reproductive cycle (Cristol & Johnsen 1994; Torkarz et al. 1998; Woodley & Moore 1999; Thurin & Aron 2008), food availability (Nicieza & Metcalfe 1997; Downs & Ratnieks 2000; Kim 2000), familiarity between competitors (Fisher 1954; Ydenberg et al. 1988; Temel”