Geographical distribution of the black howler (Alouatta pigra) in Central America

By R. Horwich, Edwin D. Johnson. Primates, 2006

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Taxonomy and Distributions of Mesoamerican Primates (2006, background): “with vellerosus, Konstant et al. (1985) were doubtful of its validity, and SilvaLópez et al.”
Population Size and Characteristics of Alouatta pigra Before and After a Major Hurricane (2007, background): “…and northern Guatemala (Crockett and Eisenberg 1987; Ostro et al. 2001; Estrada et al. 2002; Rylands et al. 2006) in a variety of habitats (Crockett 1998; Silver et al. 1998; Estrada et al. 2004), including riverine and seasonally flooded areas (Horwich and Johnson 1986; Silver et al. 1998).”
Alpha male black howler monkey responses to loud calls: effect of numeric odds, male companion behaviour and reproductive investment (2004, background): “Black howler monkeys are found in Belize, eastern Guatemala and southern Mexico (Horwich & Johnson 1986).”