Nickel defends the South African hyperaccumulator Senecio coronatus (Asteraceae) against Helix aspersa (Mollusca: Pulmonidae)

By R. Boyd, M. A. Davis, M. Wall, K. Balkwill. Chemoecology, 2002

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Heavy metal hyperaccumulating plants: how and why do they do it? And what makes them so interesting? (2011, background): “caerulescens [132] and low-Ni Senecio coronatus [133] when offered a choice between plants containing either low or high-metal concentrations.”
Environmental cleanup using plants: biotechnological advances and ecological considerations (2006, background): “Nickel or zinc hyperaccumulating plants have been shown to be protected against herbivory, due to the toxicity of the metal in their tissues (Boyd et al. 1994, 2002; Pollard and Baker 1997).”
The defense hypothesis of elemental hyperaccumulation: status, challenges and new directions (2007, background): “Deterrence of herbivores has been demonstrated for all elements studied: As (Rathinasabapathi et al. 2007), Cd (Scheirs et al. 2006), Ni (Boyd et al. 2002; Boyd and Martens 1994), Se (Hanson et al. 2004), and Zn (Pollard and Baker 1997).”