Cooperative problem solving in a social carnivore

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A comparison of innovative problem-solving abilities between wild and captive spotted hyaenas, Crocuta crocuta (2013, methodology): “Previous studies found no effect of these treatments on communication, cognition or social behaviour (Drea et al. 2002; Drea & Carter 2009).”
Greetings promote cooperation and reinforce social bonds among spotted hyaenas (2011, background): “Hyaenas are highly cooperative (Holekamp et al. 2007; Drea & Carter 2009), so if this hypothesis is correct, then greetings might promote coalition formation among adult female hyaenas during group hunting, territory defence or intragroup disputes.”
A Framework to Describe, Analyze and Generate Interactive Motor Behaviors (2012, background): “2 illustrates an interactive task that has been used to examine the social behavior of animals such as chimpanzes [36], elephants [37] and hyenas [38].”