High dissolved extracellular enzymatic activity in the deep central Atlantic ocean

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Bacterial Extracellular Enzymatic Activity in Globally Changing Aquatic Ecosystems (2010, background): “Extracellular enzymes are proportionally more important in the decomposition of particulate or colloidal material in the dark ocean [17].”
Extracellular enzymes in terrestrial, freshwater, and marine environments: perspectives on system variability and common research needs (2013, background): “Consequently, with few exceptions (e.g. Baltar et al. 2009, 2010, 2013), most investigations of enzyme activities in marine waters have been carried out in surface- and nearsurface waters (upper ca. 200 m of the ocean’s average 4,000 m depth) or in shallow coastal zones.”
Mechanisms of Microbial Carbon Sequestration in the Ocean - Future Research Directions (2014, background): “The presence of eddies has also been related to increased bacterial abundance (Arístegui and Montero, 2005) and production (Bode et al., 2001; Baltar et al., 2007), even in the mesopelagic zone (Baltar et al., 2010a).”
Emerging concepts on microbial processes in the bathypelagic ocean-ecology, biogeochemistry, and genomics (2010, background): “In the central Atlantic deep waters, Baltar et al. (2010) found high extracellular enzymatic activities in the dissolved fraction, leading to the hypothesis that microbes associated with aggregates released large amounts of free enzymes.”