Productivity increases with variation in aggression among group members in Temnothorax ants

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Personality in social insects: How does worker personality determine colony personality? (2012, background): “Population density, another environmental feature, is related to the aggression level of T. longispinosus colonies (Modlmeier and Foitzik, 2011).”
The keystone individual concept: an ecological and evolutionary overview (2014, background): “A limited but growing number of empirical tests have recovered support for this theory’s predictions, as groups that contain a diverse mixture of behavioural types tend to outperform monotypic groups (Modlmeier & Foitzik, 2011; Modlmeier, Liebmann, & Foitzik, 2012; Pruitt & Riechert, 2011).”
The development of collective personality: the ontogenetic drivers of behavioral variation across groups (2014, background): “In the closely related species, T. longispinosus, colonies also exhibit higher levels of aggression and productivity as the population becomes more crowded (Modlmeier and Foitzik, 2011).”
Linking levels of personality: personalities of the ‘average’ and ‘most extreme’ group members predict colony-level personality (2013, background): “Such elements of group composition can have large impacts on colony-level life history, productivity and reproductive output (Modlmeier & Foitzik 2011; Modlmeier et al. 2012; Pruitt 2012), and may themselves be important targets of selection.”