What determines conformity to Bergmann's rule?

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The geography of body size - challenges of the interspecific approach (2007, background): “Attempts to find which species tend to show size clines, however, have found only weak and idiosyncratic correlates for this tendency (e.g. Ashton, 2004, Meiri et al. , 2004, 2007).”
The mating system of the brown bear Ursus arctos (2012, background): “Body size varies geographically and seasonally, and is related to local food supply (Hilderbrandt et al. 1999, Meiri et al. 2007, Garshelis 2009).”
Regulation of animal size by eNPP, Bergmann's rule, and related phenomena (2011, background): “It is well known that the smallest bears occur in the tropics (Helarctos malayanus) and the largest bears inhabit temperate latitudes and subarctic and arctic areas with high availability of food from marine sources (Rausch 1963, Nowak and Paradiso 1983, Welch et al. 1997, Meiri et al. 2007).”
Bergmann's rule in mammals: a cross-species interspecific pattern (2013, unknown): “(Meiri et al. 2007), or phylogeny (de Queiroz and Ashton 2004).”