Feather mites on birds: costs of parasitism or conditional outcomes?

By G. Blanco, J. Tella, J. Potti, A. Baz. 2001

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The evolutionary transition to coloniality promotes higher blood parasitism in birds (2002, background): “Feather mites are benign ectosymbionts transmitted by host-to-host body contact (Blanco et al. 2001; Jovani et al., 2001b), and the number of congeners that directly contact with an individual may be higher in winter ¯ocks and communal roosts than in the nests, as transmission at the nests would be…”
How Birds Combat Ectoparasites (2010, background): “[102] suggested that feather mites are commensals, or even mutualists, rather than parasites.”