Can vulnerability among British bumblebee (Bombus) species be explained by niche position and breadth

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The same, but different: pollen foraging in honeybee and bumblebee colonies (2011, background): “2008), climatic niche (Williams et al. 2007; 2009), emergence time of queens (Fitzpatrick et al.”
Climatic Risk and Distribution Atlas of European Bumblebees (2015, background): “Species at their range margins can be particularly sensitive to additional threats apart from climatic limitations (Williams, 1988; Thomas et al., 2001; Williams et al., 2007; Oliver et al., 2009).”
Evidence for decline in eastern North American bumblebees (Hymenoptera: Apidae), with special focus on Bombus affinis Cresson (2008, background): “These results support the suggestion by Williams et al. (2007) that the causes of bumblebee decline likely differ between Europe and North America.”
Species' intrinsic traits inform their range limitations and vulnerability under environmental change (2015, unknown): “Species ranges are also determined by their environmental tolerances, for example the range of climatic conditions and the diversity of vegetation types (‘habitats’) they exploit (e.g. Thompson et al., 1999; Williams et al., 2007).”
The bumblebees of Sichuan (Hymenoptera: Apidae, Bombini) (2009, background): “These changes do not seem to be driven by climate change (Williams, 1986; Williams et al., 2007).”