The influence of horn and body size on the reproductive behavior of the horned rainbow scarab beetlePhanaeus difformis (Coleoptera: scarabaeidae)

By J. L. Rasmussen. Journal of insect behavior, 2005

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The behavioral ecology of threshold evolution in a polyphenic beetle (2003, background): “access to dung, which constitutes a patchy and ephemeral resource crucial for female reproduction (Hanski and Cambefort, 1991; Moczek, 1999; Otronen, 1988; Rasmussen, 1994).”
Endocrine Control of Insect Polyphenism (2012, background): “!ese male-dimorphic polyphenisms are characterized by a relatively abrupt switch between morphs that corresponds with a critical, or threshold, body size (Cook, 1987; Danforth, 1991; Diakonov, 1925; Eberhard and Gutierrez, 1991; Emlen et al., 2005a; Iguchi, 1998; Kawano, 1995; Kukuk, 1966; Rasmussen”
The Evolution of Animal Weapons (2008, background): “Both body size and horn length confer significant performance advantages to fighting males (76, 185, 198, 204, 215), and this can translate into fertilization success (124).”
Appraising Behaviour During Male-male Interaction in the Japanese Horned Beetle Trypoxylus Dichotomus Septentrionalis (Kono) (2003, background): “In several species of beetles, horn length or body size significantly affects the outcome of male-male competitions (Otronen, 1988; Rasmussen, 1994; Emlen, 1997a; Moczek & Emlen, 2000).”