How do marine diatoms fix 10 billion tonnes of inorganic carbon per year

By E. Granum, J. Raven, R. Leegood. 2005

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Advances in understanding the cyanobacterial CO2-concentrating-mechanism (CCM): functional components, Ci transporters, diversity, genetic regulation and prospects for engineering into plants. (2008, background): “, 2004) and diatoms (Granum et al., 2005) that allow the C4 cycle to proceed in a single cell, but with a level of intracellular compartmentalization of biochemical reactions].”
Acclimation to elevated carbon dioxide and ultraviolet radiation in the diatom Thalassiosira pseudonana: Effects on growth, photosynthesis, and spectral sensitivity of photoinhibition (2008, background): “Key enzymes of the C4 photosynthetic pathway occur in its genome, suggesting that a C4-based CCM can operate in some diatoms (Reinfelder et al. 2000), but many aspects still need clarification (Armbrust et al. 2004; Granum et al. 2005).”
The effect of pCO2 on carbon acquisition and intracellular assimilation in four marine diatoms (2009, methodology): “Also controversially discussed in diatoms is the potential role of a C 4 like photosynthetic pathway within carbon assimilation (Reinfelder et al., 2000, 2004; Granum et al., 2005; Roberts et al., 2007a,b; Kroth et al., 2008).”