An Overview of the Classification and Evolution of the Major Scarab Beetle Clades (Coleoptera: Scarabaeoidea) Based on Preliminary Molecular Analyses

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The beetle tree of life reveals that Coleoptera survived end‐Permian mass extinction to diversify during the Cretaceous terrestrial revolution (2015, background): “The strongly supported clade Lucanidae+Nicaginae+Diphyllostomatidae (BPP= 1.0) was also found by Smith et al. (2006), whereas Lucanidae+Glaresidae were sister groups in Ahrens et al. (2014).”
Phylogeny and evolution of Staphyliniformia and Scarabaeiformia: forest litter as a stepping stone for diversification of nonphytophagous beetles (2015, methodology): “For example, Smith et al. (2006), using only parsimony analyses, found a weakly supported clade consisting of Geotrupidae (including Bolboceratinae), Pleocomidae and Passalidae.”
A Review of the Family-group Names for the Superfamily Scarabaeoidea (Coleoptera) with Corrections to Nomenclature and a Current Classification (2006, methodology): “Therefore, I decided to take a somewhat conservative approach for each with some consideration given to molecular phylogenetics results (Smith et al. 2006).”