Multimodal signalling: structural ultraviolet reflectance predicts male mating success better than pheromones in the butterfly Colias eurytheme L. (Pieridae)

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Quantifying iridescent coloration in animals: a method for improving repeatability (2011, methodology): “…variation in butterfly iridescent coloration (e.g., Kemp 2006a, b, 2008b; Kemp and Macedonia 2006; Kemp et al. 2006; Kemp and Rutowski 2007; Papke et al. 2007; Rutowski et al. 2010), but has only been utilized in one study on an iridescent bird (Madsen et al. 2007), and to our knowledge,…”
Female mating biases for bright ultraviolet iridescence in the butterfly Eurema hecabe (Pieridae) (2008, background): “Because body size may offer a phenotypic marker of mate quality in both male and female butterflies and because both sexes may be choosy (Rutowski 1985), I incorporated this parameter along with UV reflectance in the assessment of mating patterns and potential mechanisms of male and female mate choi”