Variation in abundance of foxes (Vulpes vulpes) between three regions of rural Britain, in relation to landscape and other variables

By M. Heydon, J. Reynolds, M. Short. 2000

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A review of methods to estimate the abundance of terrestrial carnivores using field signs and observation (2001, methodology): “1991), and to calculate densities (Heydon et al. 2000) using the distance-sampling method (Buckland et al.”
Surveying carnivores at large spatial scales: a comparison of four broad-applied methods (2007, background): “…are usually scarce, elusive and nocturnal, they are very difficult to count by traditional methods such as transects based on direct observation (e.g. spotlight or line-transects, Heydon et al. 2000; Sharp et al. 2001) or capture-recapture estimates based on trapping records (Tuyttens et al. 1999).”
Winter severity limits red fox populations in Eurasia (2007, background): “Winter mortality may limit fox populations, and indeed some studies have shown that high mortality in red foxes, caused by, for example, intense culling by gamekeepers, can suppress fox populations (Heydon et al., 2000).”