Adult–Adult Play in Primates: Comparative Analyses of its Origin, Distribution and Evolution

By S. Pellis, A. Iwaniuk. 2000

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Sharing the motivation to play: the use of signals in adult bonobos (2008, background): “Some studies (Pellis et al. 1993; Pellis & Iwaniuk 2000; Palagi 2006) emphasized that contact play (in its more intense variants, e.g.”
A comparison of bonobo and chimpanzee tool use: evidence for a female bias in the Pan lineage (2010, unknown): “In chimpanzees, however, adult play is very rarely observed (Pellis & Iwaniuk 2000 ).”
Social tolerance and adult play in macaque societies: a comparison with different human cultures (2012, background): “…inmany species social play continues into adulthood (ungulates: Aldis 1975; rodents: Pellis 2002; canids: Bauer & Smuts 2007; nonhuman primates: Pellis & Iwaniuk 2000; humans: Power 2000), thus suggesting that this behaviour can provide benefits throughout the postbirth life cycle (Palagi et…”
Short-Term Benefits of Play Behavior and Conflict Prevention in Pan paniscus (2006, unknown): “…motor and cognitive skills (Dolhinow, 1999; Fagen, 1981; Loizos, 1967; Martin and Caro, 1985; Poirier et al., 1978), improving individual fighting skills, performing social assessment (Pellis and Iwaniuk, 1999, 2000; Thompson, 1998), and training for the unexpected (Špinka et al., 2001).”